Principles of Biological Oceanography

Faculteit Science and Engineering
Jaar 2021/22
Vakcode WMMB003-05
Vaknaam Principles of Biological Oceanography
Niveau(s) master
Voertaal Engels
Periode semester I a (06-09-2021 till 26-09-2021)

Uitgebreide vaknaam Principles of Biological Oceanography
Leerdoelen At the end of the course, the student is able to:

1. Identify and describe marine pelagic organisms and habitats, including phytoplankton, zooplankton and nekton.

2. Explain and/or interpret the fundamental physical, chemical and ecological factors that drive structure and functioning of pelagic food-webs.

3. Identify current topics in Biological Oceanography and explain how pelagic communities are affected by human impacts including climate change.

4. Identify phyto-, zooplankton and nekton from the class to species levels.

5. Learn basic marine sampling and analyses (e.g. water sampling, CTD profiling, calculating biomass, cell densities)
Omschrijving The course includes classroom lectures, practicals and a 2-day excursion to NIOZ. First, a series of introductory lectures will be given, starting with basic physical and chemical oceanography, followed by phytoplankton, zooplankton, and nekton diversity and productivity. Emphasis will be given to bottom-up controls of marine pelagic functioning and productivity. The second series of lectures is dedicated to Current Topics in Biological Oceanography, related to human impacts, including global climate change. These may encompass topics like oligotrophication, harmful algal blooms, pollution, and fisheries. Practicals focus on the recognition of major taxonomic groups and species. During the two days excursion, the variety of NIOZ research at both Texel and Yerseke locations will be demonstrated.
Uren per week
Onderwijsvorm Hoorcollege (LC), Practisch werk (PRC)
(Practicals and excursion need to be attended and passed)
Toetsvorm Practisch werk (PR), Schriftelijk tentamen (WE)
(Final mark is based on the written exam. Practicals and excursion need to be attended and passed. However students do not receive a final mark until a missed practical is compensated.)
Vaksoort master
Coördinator dr. W.H. van de Poll
Docent(en) dr. T. Gerkema ,dr. M. Middag ,dr. E.J. Stamhuis ,prof. dr. K.R. Timmermans ,dr. W.H. van de Poll
Verplichte literatuur
Titel Auteur ISBN Prijs
Powerpoints of lectures and additional reading material (review articles, not mandatory for examination but can be used as supporting material). Manual for the practicals.
Entreevoorwaarden The course unit assumes prior knowledge acquired from a bachelor degree in Biology.
Opmerkingen Register for this course by sending an email to:

Lecture 1 22 hours
Practical 1 24 hours
Practical 2 12 hours
Self-study 85 hours
Total 140 hours
Locations: Linneausborg, Nijenborgh 7, Groningen;
NIOZ, ‘t Horntje, Texel, NIOZ, Yerseke.
Opgenomen in
Opleiding Jaar Periode Type
MSc Ecology and Evolution: Ecology and Conservation  (Electives/optional modules) - semester I a keuze
MSc Ecology and Evolution: Evolutionary Biology  (Electives/optional modules) - semester I a keuze
MSc Marine Biology: Research  ( Track Research) - semester I a verplicht
MSc Marine Biology: Science, Business and Policy - semester I a verplicht