Hemelrijk, Prof. Charlotte
Charlotte Hemelrijk is professor of self-organization in social systems. Within the department of Behavioural ecology and self-organisation, she concentrates on primates, fish and birds. Hemelrijk studied biology and gained her PhD with a thesis on the social behaviour of chimpansees. From 1991 to 2003, she worked at the laboratory of Artificial Intelligence at the University of Zurich, where she conducted research on artificial social systems and she obtained her habilitation.
In 2003 she became a Rosalind Franklin Fellow at the University of Groningen, where she was appointed Professor of Self-organization of Social Systems in 2006. Hemelrijk and her department use computer models to examine the extent to which complex natural phenomena are created by self-organization, as a side effect of the interaction between individuals and their social and physical environment. Her fields of research include starling flocks and fish schools.
Video
- The mystery of massive swarms (Hemelrijk from 4.20)
Previously in the news
Contact and further information
- More about Hemelrijk (with flock of starlings video)
- See profile page
Publications
2022
Hemelrijk,
C. K., Seex, L., Pederboni, M., Ilany, A.,
Geffen, E., & Koren, L. (2022). Adult sex ratios and
partial dominance of females over males in the rock hyrax.
Frontiers in Ecology and Evolution,
10, [1004919]. https://doi.org/10.3389/fevo.2022.1004919
Storms,
R. F., Carere, C., Musters, R., van Gasteren,
H., Verhulst, S., & Hemelrijk, C. K.
(2022). Deterrence of birds with an artificial predator, the
RobotFalcon. Journal of the Royal Society
Interface, 19(195), 20220497. https://doi.org/10.1098/rsif.2022.0497
Papadopoulou,
M., Hildenbrandt, H., & Hemelrijk, C.
K. (2022). Disentangling patterns of collective
escape in starling flocks. Poster session presented at
European Conference on Behavioural Biology 2022, Groningen,
Netherlands.
Costanzo,
A., Haeringen ,van, E., & Hemelrijk, C.
(2022). Effect of time-delayed interactions on milling: A
minimal model. EPL (Europhysics
Letters), 138(2), [22002]. https://doi.org/10.1209/0295-5075/ac5ed1
Papadopoulou,
M., Hildenbrandt, H., Sankey, D. W. E.,
Portugal, S. J., & Hemelrijk, C. K. (2022).
Emergence of splits and collective turns in pigeon flocks
under predation. Royal Society Open
Science, 9(2), [211898]. https://doi.org/10.1098/rsos.211898
Testori,
M., Hemelrijk, C. K., & Beersma, B. (2022).
Gossip promotes cooperation only when it is pro-socially
motivated. Scientific Reports,
12(1), 4790. https://doi.org/10.1038/s41598-022-08670-7
van
Haeringen, E., & Hemelrijk, C. (2022).
Hierarchical development of dominance through the
winner-loser effect and socio-spatial structure.
PLoS ONE, 17(2 February),
[e0243877]. https://doi.org/10.1371/journal.pone.0243877
Seex,
L., Saccà, T., & Hemelrijk, C.
K. (2022). How to measure intersexual
dominance? Frontiers in Ecology and
Evolution. https://doi.org/10.3389/fevo.2022.982507
Seex,
L., Fichtel, C., Kappeler, P. M., & Hemelrijk, C.
K. (2022). Interrelationship among spatial cohesion,
aggression rate, counter-aggression and female dominance in three
lemur species. Behavioral Ecology and
Sociobiology, 76(10), [133]. https://doi.org/10.1007/s00265-022-03241-2
Saccá,
T., van de Waal, E., & Hemelrijk, C. K.
(2022). Male intrasexual aggression and partial dominance
of females over males in vervet monkeys. Poster session
presented at European Conference on Behavioural Biology 2022,
Groningen, Netherlands.
Saccà,
T., Gort, G., van de Waal, E., & Hemelrijk, C. K.
(2022). Male intrasexual aggression and partial dominance of
females over males in vervet monkeys. Frontiers in
Ecology and Evolution, 10, [930266]. https://doi.org/10.3389/fevo.2022.930266
Storms,
R., Carere, C., Verhulst, S., &
Hemelrijk, C. K. (2022). Patterns of collective
escape of bird flocks under predation by a RobotFalcon.
Poster session presented at European Conference on Behavioural
Biology 2022, Groningen, Netherlands.
Saccá,
T., Gort, G., van de Waal, E., & Hemelrijk, C.
K. (2022). Reducing the bias due to unknown
relationships in measuring the steepness of a dominance
hierarchy. Poster session presented at European
Conference on Behavioural Biology 2022, Groningen,
Netherlands.
Papadopoulou,
M., Hildenbrandt, H., Sankey, D. W. E.,
Portugal, S. J., & Hemelrijk, C. K. (2022).
Self-organization of collective escape in pigeon
flocks. PLoS Computational Biology,
18(1), [e1009772]. https://doi.org/10.1371/journal.pcbi.1009772
Kappeler,
P. M., Huchard, E., Baniel, A., Canteloup, C., Charpentier, M. J.
E., Cheng, L., Davidian, E., Duboscq, J., Fichtel, C.,
Hemelrijk, C. K., Höner, O. P., Koren, L., Micheletta,
J., Prox, L., Saccà, T., Seex, L., Smit, N.,
Surbeck, M., van de Waal, E., & Girard-Buttoz, C. (2022).
Sex and dominance: How to assess and interpret intersexual
dominance relationships in mammalian societies.
Frontiers in Ecology and Evolution,
10, [918773]. https://doi.org/10.3389/fevo.2022.918773
2021
Costanzo,
A., Hildenbrandt, H., & Hemelrijk, C.
K. (2022). Causes of variation of darkness in flocks
of starlings, a computational model. Swarm
intelligence, 16, 91-105. https://doi.org/10.1007/s11721-021-00207-4
Sankey,
D. W. E., Storms, R. F., Musters, R. J.,
Russell, T. W., Hemelrijk, C. K., & Portugal, S.
J. (2021). Absence of “selfish herd” dynamics in
bird flocks under threat. Current
Biology, 31(14), 3192-3198.e7. https://doi.org/10.1016/j.cub.2021.05.009
Izar,
P., Fernández-Bolaños, M., Seex, L., Gort,
G., Suscke, P., Tokuda, M., Mendonça-Furtado, O., Verderane,
M. P., & Hemelrijk, C. K. (2021). Female
emancipation in a male dominant, sexually dimorphic primate under
natural conditions. PLoS ONE,
16(4), [e0249039]. https://doi.org/10.1371/journal.pone.0249039
2020
Muller,
K., Hemelrijk, C. K., Westerweel, J., & Tam, D. S.
W. (2020). Calibration of multiple cameras for large-scale
experiments using a freely moving calibration target.
Experiments in Fluids, 61(1), [7].
https://doi.org/10.1007/s00348-019-2833-z
Hemelrijk,
C. K., Wubs, M. D., Gort, G., Botting, J., & van de
Waal, E. (2020). Corrigendum: Dynamics of intersexual
dominance and adult sex-ratio in wild vervet monkeys (Front.
Psychol., (2020), 11, (839), 10.3389/fpsyg.2020.00839).
Frontiers in Psychology, 11,
[598699]. https://doi.org/10.3389/fpsyg.2020.598699
Hemelrijk,
C. K., Wubs, M., Gort, G., Botting, J., & van de Waal,
E. (2020). Dynamics of intersexual dominance and adult
sex-ratio in wild vervet monkeys. Frontiers in
Psychology, 11, 839. [839]. https://doi.org/10.3389/fpsyg.2020.00839
van
Weerden, J. F., Verbrugge, R., &
Hemelrijk, C. K. (2020). Modelling non-attentional
visual information transmission in groups under predation.
Ecological Modelling, 431, [109073].
https://doi.org/10.1016/j.ecolmodel.2020.109073
Gompper,
G., Winkler, R. G., Speck, T., Solon, A., Nardini, C., Peruani, F.,
Löwen, H., Golestanian, R., Kaupp, U. B., Alvarez, L.,
Kiørboe, T., Lauga, E., Poon, W. C. K., DeSimone, A.,
Muiños-Landin, S., Fischer, A., Söker, N. A., Cichos, F.,
Kapral, R., ... Kale, S. (2020). The 2020 motile active
matter roadmap. Journal of Physics-Condensed
Matter, 32(19), [193001]. https://doi.org/10.1088/1361-648X/ab6348
2019
Storms,
R. F., Carere, C., Zoratto, F., & Hemelrijk, C.
K. (2019). Complex patterns of collective escape in
starling flocks under predation. Behavioral
Ecology and Sociobiology, 73(1), [10]. https://doi.org/10.1007/s00265-018-2609-0
Hemelrijk,
C. K., Costanzo, A., Hildenbrandt,
H., & Carere, C. (2019). Damping of waves of
agitation in starling flocks. Behavioral Ecology
and Sociobiology, 73(9), [125]. https://doi.org/10.1007/s00265-019-2734-4
Mills,
R., Taylor, G. K., & Hemelrijk, C. K.
(2019). Sexual size dimorphism, prey morphology and catch
success in relation to flight mechanics in the peregrine falcon: A
simulation study. Journal of Avian
Biology, 50(3), [01979]. https://doi.org/10.1111/jav.01979
2018
Mills,
R., Hildenbrandt, H., Taylor, G. K.,
& Hemelrijk, C. K. (2018). Physics-based
simulations of aerial attacks by peregrine falcons reveal that
stooping at high speed maximizes catch success against agile
prey. PLoS Computational Biology,
14(4), [1006044]. https://doi.org/10.1371/journal.pcbi.1006044
Lecheval,
V., Jiang, L., Tichit, P., Sire, C., Hemelrijk, C. K.,
& Theraulaz, G. (2018). Social conformity and propagation
of information in collective U-turns of fish schools.
Proceedings of the Royal Society of London. Series B,
Biological Sciences, 285(1877), [20180251]. https://doi.org/10.1098/rspb.2018.0251
Costanzo,
A., & Hemelrijk, C. K. (2018).
Spontaneous emergence of milling (vortex state) in a
Vicsek-like model. Journal of Physics D: Applied
Physics, 51(13), [134004]. https://doi.org/10.1088/1361-6463/aab0d4
Tan,
A. W. Y., Hemelrijk, C. K., Malaivijitnond, S., &
Gumert, M. D. (2018). Young macaques (Macaca
fascicularis) preferentially bias attention towards closer,
older, and better tool users. Animal
Cognition, 21(4), 551-563. https://doi.org/10.1007/s10071-018-1188-9
2017
Jager,
W., Verbrugge, R., Flache,
A., de Roo, G., Hoogduin,
L., & Hemelrijk, C. (Eds.) (2017).
Advances in Social Simulation 2015. (Advances
in Intelligent Systems and Computing; Vol. 528). Springer
International Publishing. https://doi.org/10.1007/978-3-319-47253-9
Jager,
W., Verbrugge, R., Flache,
A., de Roo, G., Hoogduin,
L., & Hemelrijk, C. (2017).
Introduction. In W. Jager, R. Verbrugge, A. Flache, G.
de Roo, L. Hoogduin, & C. Hemelrijk (Eds.), Advances in
Social Simulation 2015 (pp. I-XVIII). (Advances in Intelligent
Systems and Computing; Vol. 528). Springer.
Hemelrijk,
C. (2017). Simulating complexity of animal social
behaviour. In B. Edmonds, & R. Meyer (Eds.),
Simulating Social Complexity : A Handbook (pp. 581-616).
(Understanding Complex Systems). Springer Verlag. https://doi.org/10.1007/978-3-540-93813-2_22
Hogan,
B. G., Hildenbrandt, H., Scott-Samuel, N. E., Cuthill,
I. C., & Hemelrijk, C. K. (2017). The
confusion effect when attacking simulated three-dimensional
starling flocks. Royal Society Open
Science, 4(1), [160564]. https://doi.org/10.1098/rsos.160564
Hemelrijk,
C. K., Kappeler, P. M., & Puga-Gonzalez, I. (2017).
The self-organization of social complexity in group-living
animals: Lessons from the DomWorld model. In M. Naguib, J.
Podos, LW. Simmons, L. Barrett, SD. Healy, & M. Zuk (Eds.),
Advances in the Study of Behavior (pp. 361-405). (Advances
in the Study of Behavior; Vol. 49). Academic Press. https://doi.org/10.1016/bs.asb.2017.02.005
2016
Stroebe,
K., Nijstad, B. A., & Hemelrijk, C.
K. (2017). Female dominance in human groups: Effects
of sex ratio and conflict level. Social
Psychological and Personality Science, 8(2),
209-218. https://doi.org/10.1177/1948550616664956
Kalcher-Sommersguter,
E., Preuschoft, S., Franz-Schaider, C., Hemelrijk, C.
K., Crailsheim, K., & Massen, J. J. M. (2016).
Corrigendum: Early maternal loss affects social integration
of chimpanzees throughout their lifetime.
Scientific Reports, 6, [24842]. https://doi.org/10.1038/srep24842
2015
Puga-Gonzalez,
I., Cooper, M. A., & Hemelrijk, C. K. (2016).
“Targeting or supporting, what drives patterns of
aggressive intervention in fights?”.
American Journal of Primatology,
78(2), 247-255. https://doi.org/10.1002/ajp.22505
Hemelrijk,
C., Puga-Gonzalez, I., & Steinhauser, J. (2015).
Cooperation, coalition, alliances. In W. Henke, &
I. Tattersall (Eds.), Handbook of Paleoanthropology: Primate
Evolution and Human Origins (2 ed., Vol. II, pp. 1693).
Springer.
Hemelrijk,
C. K., & Hildenbrandt, H. (2015).
Diffusion and topological neighbours in flocks of starlings:
relating a model to empirical data. PLoS
ONE, 10(5), [e0126913]. https://doi.org/10.1371/journal.pone.0126913
Kalcher-Sommersguter,
E., Preuschoft, S., Franz-Schaider, C., Hemelrijk, C.
K., Crailsheim, K., & Massen, J. J. M. (2015).
Early maternal loss affects social integration of chimpanzees
throughout their lifetime. Scientific
Reports, 5, [16439]. https://doi.org/10.1038/srep16439
Puga-Gonzalez,
I., Hoscheid, A., & Hemelrijk, C. K. (2015).
Friendship, reciprocation, and interchange in an
individual-based model. Behavioral Ecology and
Sociobiology, 69(3), 383-394. https://doi.org/10.1007/s00265-014-1850-4
Demsar,
J., Hemelrijk, C. K., Hildenbrandt, H.,
& Bajec, I. L. (2015). Simulating predator attacks on
schools: Evolving composite tactics. Ecological
Modelling, 304, 22-33. https://doi.org/10.1016/j.ecolmodel.2015.02.018
van
der Post, D. J., Verbrugge, R., & Hemelrijk,
C. K. (2015). The evolution of different forms of
sociality: Behavioral mechanisms and eco-evolutionary
feedback. PLoS ONE, 10(1),
[e0117027]. https://doi.org/10.1371/journal.pone.0117027
Hemelrijk,
C. (Photographer). (2015). Waves of terror in Starling
flocks: Specific reactions of Starling flocks to predator
attacks. Web publication/site, British Ornithologists'
Union. https://www.bou.org.uk/waves-of-terror-in-starling-flocks/
Hemelrijk,
C. K., van Zuidam, L., & Hildenbrandt, H.
(2015). What underlies waves of agitation in starling
flocks. Behavioral Ecology and
Sociobiology, 69(5), 755-764. https://doi.org/10.1007/s00265-015-1891-3
2014
Hemelrijk,
C. K., & Hildenbrandt, H. (2015).
Scale-free correlations, influential neighbours and speed
control in flocks of birds. Journal of Statistical
Physics, 158(3), 563-578. https://doi.org/10.1007/s10955-014-1154-0
Hemelrijk,
C. K., Reid, D. A. P., Hildenbrandt, H., &
Padding, J. T. (2015). The increased efficiency of fish
swimming in a school. Fish and
Fisheries, 16(3), 511-521. https://doi.org/10.1111/faf.12072
Puga-Gonzalez,
I., Butovskaya, M., Thierry, B., & Hemelrijk, C.
K. (2014). Empathy versus parsimony in understanding
post-conflict affiliation in monkeys: Model and empirical
data. PLoS ONE, 9(3),
[e91262]. https://doi.org/10.1371/journal.pone.0091262
van
der Vaart, E., & Hemelrijk, C. K. (2014).
'Theory of Mind' in animals: Ways to make progress.
Synthese, 191(3), 335-354. https://doi.org/10.1007/s11229-012-0170-3
2013
van
Weerden, J. F., Reid, D. A. P., & Hemelrijk, C.
K. (2014). A meta-analysis of steady undulatory
swimming. Fish and Fisheries,
15(3), 397-409. https://doi.org/10.1111/faf.12022
Krueger,
K., Flauger, B., Farmer, K., & Hemelrijk, C.
(2014). Movement initiation in groups of feral horses.
Behavioural Processes, 103, 91-101.
https://doi.org/10.1016/j.beproc.2013.10.007
van
der Post, D. J., de Weerd, H., Verbrugge,
R., & Hemelrijk, C. K. (2013). A
novel mechanism for a survival advantage of vigilant individuals in
groups. American Naturalist,
182(5), 682-688. https://doi.org/10.1086/673298
Schroeder,
J., Dugdale, H. L., Radersma, R., Hinsch, M., Buehler,
D. M., Saul, J., Porter, L., Liker, A., De Cauwer, I., Johnson, P.
J., Santure, A. W., Griffin, A. S., Bolund, E., Ross, L., Webb, T.
J., Feulner, P. G. D., Winney, I., Szulkin, M., Komdeur,
J., ... Horrocks, N. (2013). Fewer invited talks by
women in evolutionary biology symposia. Journal of
Evolutionary Biology, 26(9), 2063-2069. https://doi.org/10.1111/jeb.12198
2012
Hemelrijk,
C. K., & Puga-Gonzalez, I. (2012). An
individual-oriented model on the emergence of support in fights,
its reciprocation and exchange. PLoS
ONE, 7(5), [e37271]. https://doi.org/10.1371/journal.pone.0037271
van
der Vaart, E., Hemelrijk, C., & Verbrugge,
R. (2012). Corvid re-chaching without 'Theory of
Mind': A model. PLoS ONE, 7,
[e32904]. https://doi.org/10.1371/journal.pone.0032904
Reid,
D. A. P., Hildenbrandt, H., Hemelrijk, C.
K., & Padding, J. T. (2012). Fluid dynamics of
moving fish in a two-dimensional multiparticle collision dynamics
model. Physical Review E,
85(2), [021901]. https://doi.org/10.1103/PhysRevE.85.021901
Hemelrijk,
C. K., & Hildenbrandt, H. (2012).
Schools of fish and flocks of birds: Their shape and internal
structure by self-organization. Interface
Focus, 2(6), 726-737. https://doi.org/10.1098/rsfs.2012.0025
Kunz,
H., & Hemelrijk, C. K. (2012). Simulations
of the social organization of large schools of fish whose
perception is obstructed. Applied Animal Behaviour
Science, 138(3-4), 142-151. https://doi.org/10.1016/j.applanim.2012.02.002
2011
Hemelrijk,
C. K., & Bolhuis, J. J. (2011). A minimalist
approach to comparative psychology. Trends in
Cognitive Sciences, 15(5), 185-186. https://doi.org/10.1016/j.tics.2011.02.007
van
der Vaart, E., Verbrugge, R., & Hemelrijk,
C. K. (2011). Corvid caching: Insights from a
cognitive model. Journal of Experimental
Psychology: Animal Behavior Processes, 37(3),
330-340. https://doi.org/10.1037/a0022988
Procaccini,
A., Orlandi, A., Cavagna, A., Giardina, I., Zoratto, F., Santucci,
D., Chiarotti, F., Hemelrijk, C. K., Alleva, E.,
Parisi, G., & Carere, C. (2011). Propagating waves in
starling, Sturnus vulgaris, flocks under predation.
Animal Behavior, 82(4), 759-765. https://doi.org/10.1016/j.anbehav.2011.07.006
Hemelrijk,
C. K. (2011). Simple reactions to nearby neighbors and
complex social behavior in primates. In R. Menzel, & J.
Fischer (Eds.), Animal Thinking: Contemporary Issues in
Comparative Cognition (pp. 223-238). The MIT Press.
Hemelrijk,
C. (2011). Simple reactions to nearby neighbours and
complex social behaviour in primates. In R. Menzel, & J.
Fischer (Eds.), Animal Thinking: Contemporary Issues in
Comparative Cognition (Vol. 8, pp. 223-238). MIT Press. http://www.jstor.org/stable/j.ctt5vjs6z.19
Hemelrijk,
C. K., Jensen, K., Silk, J. B., Andrews, K., Bshary, R.,
& Cheney, D. L. (2011). Social knowledge. In R.
Menzel, & J. Fischer (Eds.), Animal Thinking: Contemporary
Issues in Comparative Cognition (Vol. 8, pp. 267-292). http://www.jstor.org/stable/j.ctt5vjs6z.22
Hemelrijk,
C. K., & Hildenbrandt, H. (2011).
Some causes of the variable shape of flocks of birds.
PLoS ONE, 6(8), [22479]. https://doi.org/10.1371/journal.pone.0022479
2010
Oosten,
J. E., Magnhagen, C., & Hemelrijk, C. K. (2010).
Boldness by habituation and social interactions: A
model. Behavioral Ecology and
Sociobiology, 64(5), 793-802. https://doi.org/10.1007/s00265-009-0896-1
Hemelrijk,
C. K., Hildenbrandt, H., Reinders, J.,
& Stamhuis, E. J. (2010). Emergence of oblong
school shape: Models and empirical data of fish.
Ethology, 116(11), 1099-1112. https://doi.org/10.1111/j.1439-0310.2010.01818.x
Dijkstra,
P., Lindström, J., Metcalfe, N. B., Hemelrijk, C.
K., Brendel, M., Seehausen, O., & Groothuis, T. G.
G. (2010). Frequency-dependent social dominance in a
color polymorphic cichlid fish.
Evolution, 64(10), 2797-2807. https://doi.org/10.1111/j.1558-5646.2010.01046.x
Hildenbrandt,
H., Carere, C., & Hemelrijk, C. K. (2010).
Self-organized aerial displays of thousands of starlings: A
model. Behavioral Ecology,
21(6), 1349-1359. https://doi.org/10.1093/beheco/arq149
2009
Becher,
M. A., Hildenbrandt, H., Hemelrijk, C.
K., & Moritz, R. F. A. (2010). Brood temperature,
task division and colony survival in honeybees: A model.
Ecological Modelling, 221(5),
769-776. https://doi.org/10.1016/j.ecolmodel.2009.11.016
Puga-Gonzalez,
I., Hildenbrandt, H., & Hemelrijk, C.
K. (2009). Emergent patterns of social affiliation in
primates: A model. PLoS Computational
Biology, 5(12), [e1000630]. https://doi.org/10.1371/journal.pcbi.1000630
Reid,
D. A. P., Hildenbrandt, H., Padding, J. T.,
& Hemelrijk, C. K. (2009). Flow around fishlike
shapes studied using multiparticle collision dynamics.
Physical Review E, 79(4), [046313].
https://doi.org/10.1103/PhysRevE.79.046313
van
der Vaart, E. E., Hemelrijk, C., &
Verbrugge, L. (2009). Learning where (not) to cache: A
cognitive model for corvids. In N. A. Taatgen, & R. van
Rijn (Eds.), Proceedings of the 31st Annual Meeting of the
Cognitive Science Society (pp. 2420-2425). Cognitive Science
Society.
van
der Vaart, E. E., Verbrugge, L., &
Hemelrijk, C. (2009). Memory Effects in the Clark's
nutcracker: A cognitive model for corvids. In A. Howes, D.
Peebles, & R. Cooper (Eds.), Proceedings of the 9th
International Conference on Cognitive Modeling (pp.
412-417).
2008
Dijkstra,
P., Hemelrijk, C., Seehausen, O., &
Groothuis, T. G. G. (2009). Color polymorphism and
intrasexual competition in assemblages of cichlid fish.
Behavioral Ecology, 20(1), 138-144.
https://doi.org/10.1093/beheco/arn125
Hemelrijk,
C. K., Wantia, J., & Isler, K. (2008). Female
dominance over males in primates: Self-organisation and sexual
dimorphism. PLoS ONE, 3(7),
[2678]. https://doi.org/10.1371/journal.pone.0002678
Hemelrijk,
C. K., & Hildenbrandt, H. (2008).
Selfâorganized shape and frontal density of fish
schools. Ethology, 114(3),
245-254. https://doi.org/10.1111/j.1439-0310.2007.01459.x
2007
Hemelrijk,
C. K., & Steinhauser, J. (2007). Cooperation,
coalition and alliances. In W. Henke, & I. Tattersal
(Eds.), Handbook of Paleoanthropology: Primate Evolution and
Human Origins (1 ed., Vol. II, pp. 1321 - 1346).
Springer.
2006
Kunz,
H., Züblin, T., & Hemelrijk, C. K. (2006).
On prey grouping an predator confusion in artificial fish
schools. In Proceedings of Tenth International
Conference on Artificial Life (pp. 365 - 371). MIT
Press.
2005
Hemelrijk,
C. K. (2005). A process-oriented approach to the
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